The results indicate that: 1) Substance P labeled terminal and part cells were distributed in the Area X; 2) Substance P labeled cells were distributed in the nucleus high vocal center (HVc), magnocellular nucleus of the anterior neostriatum (MAN), robust nucleus of the archistriatum (RA) and dorsolateral nucleus of the anterior thalamus (DLM); 3) Substance P labeled terminal and fibers were distributed in the vocal control nuclei such as nucleus dorsalis medialis (DM) and the Nucleus hypoglossi, pars tracheosyringealis (nXI-Its), and in the auditory nuclei such as the nucleus ovidalisashell (Ov shell), the shell regions of mesencephalicus lateralis, pars dorsalis (MLd shell) and the nucleus intercollicularis (ICo). 
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